1. Nolan, Victoria; Gilbert, Francis; Reed, Tom; and Reader, Tom (2022) Distribution models calibrated with independent field data predict two million ancient and veteran trees in England Ecological Applications, TBC:
  2. Crowther, Lucy I; Reader, Tom; and Gilbert, Francis (2022) The effect of agri-environment schemes on Bombus terrestris colony success Ecological Entomology, TBC:
  3. Nolan, Victoria; Gilbert, Francis; and Reader, Tom (2022) Solving sampling bias problems in presence-absence or presence-only species data using zero-inflated models Journal of Biogeography, 49: 215-232. [PDF]


  1. Bromage, Adrian; Pierce, Sarah; Reader, Tom; and Compton, Lindsey (2021) Teaching statistics to non-specialists: challenges and strategies for success Journal of Further and Higher Education, 10.1080/0309877X.2021.1879744: . [PDF]
  2. Nolan, Victoria; Reader, Tom; Gilbert, Francis; and Atikinson, Nick (2021) Historical maps confirm the accuracy of zero-inflated model predictions of ancient tree abundance in English wood-pastures Journal of Applied Ecology, 58: 2661–2672. [PDF]
  3. Leavey, Alice; Taylor, Christopher; Symonds, Matthew; Gilbert, Francis; and Reader, Tom (2021) Mapping the evolution of accurate Batesian mimicry of social wasps in hoverflies Evolution, 75: 2802-2815. [PDF]
  4. Occhiuto, Francesca; Mohallal, Eman; Gilfillan, Geoffrey D.; Lowe, Andrew; and Reader, Tom (2021) Seasonal patterns in habitat use by the harvest mouse (Micromys minutus) and other small mammals Mammalia, 85: 325-335. [PDF]
  5. D’Agostino, Daniele; Burt, John A.; Santinelli, Veronica; Vaughan, Grace O.; Fowler, Ashley M.; Reader, Tom; Taylor, Brett M.; Hoey, Andrew S.; Cavalcante, Georgenes H.; Bauman, Andrew G.; and Feary, David (2021) Growth impacts in a changing ocean: insights from two coral reef fishes in an extreme environment Coral Reefs, 40: 433-446. [PDF]


  1. Durrant, Kate; Reader, Tom; and Symonds, Matthew (2020) Pre- and post-copulatory traits working in concert: Sexual dichromatism in passerines is associated with sperm morphology Biology Letters, 16: 20190568. [PDF]
  2. D’Agostino, Daniele; Jimenez, Carlos; Reader, Tom; Hadjioannou, Louis; Heyworth, Stephanie; Aplikioti, Marilena; Argyrou, Marina; and Feary, David (2020) Behavioural traits and feeding ecology of Mediterranean lionfish and native species naiveté to lionfish predation Marine Ecology Progress Series, 638: 123-135. [PDF]
  3. Alatawi, Abdulaziz Subhi; Gilbert, Francis; and Reader, Tom (2020) Modelling terrestrial reptile species richness, distributions and habitat suitability in Saudi Arabia Journal of Arid Environments, 178: 104153. [PDF]
  4. Schaus, Jessica; Uzal, Antonio; Gentle, Louise; Baker, Philip; Bearman-Brown, Lucy; Bullion, Simone; Gazzard, Abigail; Lockwood, Hannah; North, Alexandra; Reader, Tom; Scott, Dawn; Sutherland, Chris; and Yarnell, Richard (2020) Application of the Random Encounter Model in citizen science projects to monitor animal densities Remote Sensing in Ecology and Conservation, 6: 514-528. [PDF]
  5. Nolan, Victoria; Reader, Tom; Gilbert, Francis; and Atkinson, Nick (2020) The Ancient Tree Inventory: a summary of the results of a 15 year citizen science project recording ancient, veteran and notable trees across the UK Biodiversity and Conservation, 29: 3103-3129. [PDF]
  6. Vitale, Jessica D.; Jordan, Neil R.; Gilfillan, Geoffrey D.; McNutt, J. W.; and Reader, Tom (2020) Spatial and seasonal patterns of communal latrine use by spotted hyenas (Crocuta crocuta) reflect a seasonal resource-defense strategy Behavioral Ecology and Sociobiology, 74: 120. [PDF]
  7. D’Agostino, Daniele; Burt, John A.; Reader, Tom; Vaughan, Grace O.; Chapman, B.; Santinelli, Veronica; Cavalcante, Georgenes H.; and Feary, David (2020) The influence of thermal extremes on coral reef fish behaviour in the Arabian/Persian Gulf Coral Reefs, 39: 733-744. [PDF]
  8. Bachman, Steven; Wilkin, Paul; Steven, ; Reader, Tom; Field, Richard; Weber, Odile; Nordal, Inger; and Demissew, Sebsebe (2020) Extinction risk and conservation gaps for Aloe (Asphodelaceae) in the Horn of Africa Biodiversity and Conservation, 29: 77–98. [PDF]


  1. Davison, Angus; Jackson, Hannah J; Murphy, Ellis W; and Reader, Tom (2019) Discrete or indiscrete? Redefining the colour polymorphism of the land snail Cepaea nemoralis Heredity, 123: 162–175. [PDF]
  2. Bachman, Steven; Field, Richard; Reader, Tom; Raimondo, Domitilla; Donaldson, John; Schatz, George; and Nic Lughadha, Eimear (2019) Progress, challenges and opportunities for Red Listing Biological Conservation, 234: 45-55. [PDF]
  3. Ajuria Ibarra, Helena; Kinahan, Michael; Marcetteau, Julien; Mehigan, Andrew J R; Ziegelmeier, Ross O; and Reader, Tom (2019) The significance of prey avoidance behavior for the maintenance of a predator color polymorphism Behavioral Ecology, 30: 240-248. [PDF]


  1. Taylor, Christopher H; Warrin, Jonathan; Gilbert, Francis; and Reader, Tom (2017) Which traits do observers use to distinguish Batesian mimics from their models? Behavioral Ecology, 28: 460–470. [PDF]


  1. Kettel, Esther F; Perrow, Martin R; and Reader, Tom (2016) Live-trapping in the stalk zone of tall grasses as an effective way of monitoring harvest mice (Micromys minutus) European Journal of Wildlife Research, 62: 241–245. [PDF]
  2. Morris, Rohanna L; and Reader, Tom (2016) Do crab spiders perceive Batesian mimicry in hoverflies? Behavioral Ecology, 27: 920–931. [PDF]
  3. Torre, Mónica Padilla; Briefer, Elodie F; Ochocki, Brad M; McElligott, Alan G; and Reader, Tom (2016) Mother–offspring recognition via contact calls in cattle, Bos taurus Animal Behaviour, 114: 147–154. [PDF]
  4. Noor, Dzul Azri Mohamed; Jeyapalan, Jennie N; Alhazmi, Safiah; Carr, Matthew; Squibb, Benjamin; Wallace, Claire; Tan, Christopher; Cusack, Martin; Hughes, Jaime; Reader, Tom; and others, (2016) Genome-wide methylation analysis identifies genes silenced in non-seminoma cell lines NPJ Genomic Medicine, 1: 15009. [PDF]
  5. Taylor, Christopher H; Reader, Tom; and Gilbert, Francis (2016) Hoverflies are imperfect mimics of wasp colouration Evolutionary Ecology, 30: 567–581. [PDF]
  6. Taylor, Christopher H; Reader, Tom; and Gilbert, Francis (2016) Why many Batesian mimics are inaccurate: evidence from hoverfly colour patterns Proceedings of the Royal Society B: Biological Sciences, 283: 20161585. [PDF]
  7. Pomerai, David I; Iqbal, Nooria; Lafayette, Ivan; Nagarajan, Archana; Kaviani Moghadam, Mehri; Fineberg, April; Reader, Tom; Greedy, Steve; Smartt, Chris; and Thomas, David WP (2016) Microwave fields have little effect on α-synuclein aggregation in a Caenorhabditis elegans model of Parkinson’s disease Bioelectromagnetics, 37: 116–129. [PDF]


  1. Torre, Mónica Padilla; Briefer, Elodie F; Reader, Tom; and McElligott, Alan G (2015) Acoustic analysis of cattle (Bos taurus) mother–offspring contact calls from a source–filter theory perspective Applied Animal Behaviour Science, 163: 58–68. [PDF]


  1. Ajuria Ibarra, Helena; and Reader, Tom (2014) Female-limited colour polymorphism in the crab spider Synema globosum (Araneae: Thomisidae) Biological Journal of the Linnean Society, 113: 368–383. [PDF]
  2. Edmunds, Malcolm; and Reader, Tom (2014) Evidence for Batesian mimicry in a polymorphic hoverfly Evolution, 68: 827–839. [PDF]
  3. Holland, Sara L; Reader, Tom; Dyer, Paul S; and Avery, Simon V (2014) Phenotypic heterogeneity is a selected trait in natural yeast populations subject to environmental stress Environmental Microbiology, 16: 1729–1740. [PDF]


  1. Taylor, Christopher H; Gilbert, Francis; and Reader, Tom (2013) Distance transform: a tool for the study of animal colour patterns Methods in Ecology and Evolution, 4: 771–781. [PDF]
  2. Ajuria Ibarra, H; and Reader, T (2013) Reasons to be different: do conspicuous polymorphisms in invertebrates persist because rare forms are fitter? Journal of Zoology, 290: 81–95. [PDF]


  1. Curno, Olivia; Reader, Tom; McElligott, Alan G; Behnke, Jerzy M; and Barnard, Chris J (2011) Infection before pregnancy affects immunity and response to social challenge in the next generation Philosophical Transactions of the Royal Society of London B: Biological Sciences, 366: 3364–3374. [PDF]


  1. Norfolk, Olivia; Melotte, Alice; Gilbert, Francis; Zalat, Samy; and Reader, Tom (2010) A comparative study of two agamid lizards, Laudakia stellio and Pseudotrapelus sinaitus, in southern Sinai Egyptian Journal of Biology, 12: [PDF]
  2. Newbold, Tim; Reader, Tom; El-Gabbas, Ahmed; Berg, Wiebke; Shohdi, Wael M; Zalat, Samy; El Din, Sherif Baha; and Gilbert, Francis (2010) Testing the accuracy of species distribution models using species records from a new field survey Oikos, 119: 1326–1334. [PDF]


  1. Newbold, Tim; Gilbert, Francis; Zalat, Samy; El-Gabbas, Ahmed; and Reader, Tom (2009) Climate-based models of spatial patterns of species richness in Egypt’s butterfly and mammal fauna Journal of Biogeography, 36: 2085–2095. [PDF]
  2. Curno, Olivia; Behnke, Jerzy M; McElligott, Alan G; Reader, Tom; and Barnard, Chris J (2009) Mothers produce less aggressive sons with altered immunity when there is a threat of disease during pregnancy Proceedings of the Royal Society of London B: Biological Sciences, 276: 1047–1054. [PDF]
  3. Newbold, Tim; Reader, Tom; Zalat, Samy; El-Gabbas, Ahmed; and Gilbert, Francis (2009) Effect of characteristics of butterfly species on the accuracy of distribution models in an arid environment Biodiversity and Conservation, 18: 3629. [PDF]
  4. Dawe, Adam S; Bodhicharla, Rakesh K; Graham, Neil S; May, Sean T; Reader, Tom; Loader, Benjamin; Gregory, Andrew; Swicord, Mays; Bit-Babik, Giorgi; and Pomerai, David I (2009) Low-intensity microwave irradiation does not substantially alter gene expression in late larval and adult Caenorhabditis elegans Bioelectromagnetics, 30: 602–612. [PDF]
  5. Reader, Tom; and Duce, Ian R (2009) Intraguild interactions promote assortative mating and affect sexual attractiveness in a phytophagous fly Biological Journal of the Linnean Society, 98: 171–180. [PDF]
  6. Cambridge, Elisabeth A Lloyd (2009) REVIEWS AND BRIEF NOTICES The Quarterly Review of Biology, 84:
  7. Higginson, AD; and Reader, T (2009) Environmental heterogeneity, genotype-by-environment interactions and the reliability of sexual traits as indicators of mate quality Proceedings of the Royal Society of London B: Biological Sciences, 276: 1153–1159. [PDF]


  1. Dawe, Adam S; Nylund, Reetta; Leszczynski, Dariusz; Kuster, Niels; Reader, Tom; and De Pomerai, David I (2008) Continuous wave and simulated GSM exposure at 1.8 W/kg and 1.8 GHz do not induce hsp16-1 heat-shock gene expression in Caenorhabditis elegans Bioelectromagnetics, 29: 92–99. [PDF]
  2. Phillips, Tim; Barnard, Chris; Ferguson, Eamonn; and Reader, Tom (2008) Do humans prefer altruistic mates? Testing a link between sexual selection and altruism towards non-relatives British Journal of Psychology, 99: 555–572. [PDF]


  1. Higginson, Andrew D; Gilbert, Francis S; Reader, Tom; and Barnard, Christopher J (2007) Reduction of visitation rates by honeybees (Apis mellifera) to individual inflorescences of lavender (Lavandula stoechas) upon removal of coloured accessory bracts (Hymenoptera: Apidae) Entomologia Generalis, 165–178. [PDF]
  2. Holland, Sara; Lodwig, Emma; Sideri, Theodora; Reader, Tom; Clarke, Ian; Gkargkas, Konstantinos; Hoyle, David C; Delneri, Daniela; Oliver, Stephen G; and Avery, Simon V (2007) Application of the comprehensive set of heterozygous yeast deletion mutants to elucidate the molecular basis of cellular chromium toxicity Genome Biology, 8: R268. [PDF]


  1. Reader, Tom; Cornell, Stephen J; and Rohani, Pejman (2006) Aggregation, intraguild interactions and the coexistence of competitors on small ephemeral patches Oikos, 115: 321–333. [PDF]
  2. Reader, Tom; Higginson, Andrew D; Barnard, Christopher J; Gilbert, Francis S; and Course, Behavioural Ecology Field (2006) The effects of predation risk from crab spiders on bee foraging behavior Behavioral Ecology, 17: 933–939. [PDF]


  1. Reader, Tom; MacLeod, Ian; Elliott, Philip T; Robinson, Oliver J; and Manica, Andrea (2005) Inter-order interactions between flower-visiting insects: foraging bees avoid flowers previously visited by hoverflies Journal of Insect Behavior, 18: 51–57. [PDF]


  1. Reader, Tom (2003) Strong interactions between species of phytophagous fly: a case of intraguild kleptoparasitism Oikos, 103: 101–112. [PDF]
  2. Reader, Tom; and Hochuli, Dieter F (2003) Understanding gregariousness in a larval Lepidopteran: the roles of host plant, predation, and microclimate Ecological Entomology, 28: 729–737. [PDF]